Spanish Moss
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Spanish moss (''Tillandsia usneoides'') is an
epiphytic An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
that often grows upon large trees in tropical and subtropical climates. It is native to much of
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Bermuda ) , anthem = "God Save the King" , song_type = National song , song = " Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , e ...
, the
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the Atlantic Ocean, North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
,
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, the
Southern United States The Southern United States (sometimes Dixie, also referred to as the Southern States, the American South, the Southland, or simply the South) is a geographic and cultural region of the United States of America. It is between the Atlantic Ocean ...
, and
West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...
. It has been naturalized in
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
(
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
). It is known as "grandpa's beard" in
French Polynesia )Territorial motto: ( en, "Great Tahiti of the Golden Haze") , anthem = , song_type = Regional anthem , song = " Ia Ora 'O Tahiti Nui" , image_map = French Polynesia on the globe (French Polynesia centered).svg , map_alt = Location of Frenc ...
.Kew World Checklist of Selected Plant Families, ''Tillandsia usneoides''
/ref> Most known in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
, it commonly is found on the southern live oak (''
Quercus virginiana ''Quercus virginiana'', also known as the southern live oak, is an evergreen oak tree endemic to the Southeastern United States. Though many other species are loosely called live oak, the southern live oak is particularly iconic of the Old South. ...
'') and bald cypress (''
Taxodium distichum ''Taxodium distichum'' (bald cypress, swamp cypress; french: cyprès chauve; ''cipre'' in Louisiana) is a deciduous conifer in the family Cupressaceae. It is native to the southeastern United States. Hardy and tough, this tree adapts to a wide r ...
'') in the lowlands, swamps, and marshes of the mid-Atlantic and southeastern states, from the coast of southeastern
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
to
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
and west to southern
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
and
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. The specific name of the plant, ''usneoides'', means "resembling ''
Usnea ''Usnea'' is a genus of mostly pale grayish-green fruticose lichens that grow like leafless mini-shrubs or tassels anchored on bark or twigs.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, The genus is in the f ...
''", a lichen. While it superficially resembles its namesake, it is neither a
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
, and it is not native to
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
. It is a flowering plant (
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
) in the family
Bromeliaceae The Bromeliaceae (the bromeliads) are a family of monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain o ...
(the bromeliads) that grows hanging from tree branches in full sun through partial shade. Formerly, it was placed in the genera ''Anoplophytum'', ''Caraguata'', and ''Renealmia''. The northern limit of its natural range is Northampton County,
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
, with colonial-era reports of it in southern
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean to ...
where no populations are now known to exist. Its primary range is in the southeastern United States (including
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...
and the U.S. Virgin Islands) through
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, where the climate is warm enough and a relatively high average
humidity Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depe ...
occurs. It has been introduced to locations around the world with similar conditions, including
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
and Australia.


Description

Spanish moss consists of one or more slender stems, bearing alternate thin, curved or curly, and heavily scaled
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
long and broad, that grow vegetatively in a chain-like fashion (pendant), forming hanging structures of up to . The plant has no
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s, and its brown, green, yellow, or grey flowers are tiny and inconspicuous. It propagates both by seed and
vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
by fragments that are carried on the wind and stick to tree limbs or that are carried to other locations by birds as nesting material.


Ecology

Spanish moss is not parasitic, it is an
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
that absorbs nutrients and water through its own leaves from the air and rain falling upon it. While its presence rarely kills the trees on which it grows, it occasionally becomes so thick that, by shading the leaves of the tree, it slows the growth rate of the tree. It can use the water-conserving strategy of
crassulacean acid metabolism Crassulacean acid metabolism, also known as CAM photosynthesis, is a carbon fixation pathway that evolved in some plants as an adaptation to arid conditions that allows a plant to photosynthesize during the day, but only exchange gases at night. ...
for photosynthesis. In the southern U.S., the plant seems to show preferences for southern live oak (''
Quercus virginiana ''Quercus virginiana'', also known as the southern live oak, is an evergreen oak tree endemic to the Southeastern United States. Though many other species are loosely called live oak, the southern live oak is particularly iconic of the Old South. ...
'') and bald cypress (''
Taxodium distichum ''Taxodium distichum'' (bald cypress, swamp cypress; french: cyprès chauve; ''cipre'' in Louisiana) is a deciduous conifer in the family Cupressaceae. It is native to the southeastern United States. Hardy and tough, this tree adapts to a wide r ...
'') because of their high rates of foliar mineral leaching (calcium,
magnesium Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 of the periodic ta ...
,
potassium Potassium is the chemical element with the symbol K (from Neo-Latin ''kalium'') and atomic number19. Potassium is a silvery-white metal that is soft enough to be cut with a knife with little force. Potassium metal reacts rapidly with atmosphe ...
, and
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
) that provides an abundant supply of nutrients to the epiphytic plant. It also can colonize other tree species such as sweetgum (''
Liquidambar styraciflua American sweetgum (''Liquidambar styraciflua''), also known as American storax, hazel pine, bilsted, redgum, satin-walnut, star-leaved gum, alligatorwood, or simply sweetgum, is a deciduous tree in the genus ''Liquidambar'' native to warm temper ...
''), crepe-myrtles (''
Lagerstroemia ''Lagerstroemia'' (), commonly known as crape myrtle (also spelled crepe myrtle or crêpe myrtle), is a genus of around 50 species of deciduous and evergreen trees and shrubs native to the Indian subcontinent, southeast Asia, northern Australi ...
'' spp.), other oaks, and even
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
s. Spanish moss shelters a number of creatures, including
rat snake Rat snakes are members – along with kingsnakes, milk snakes, vine snakes and indigo snakes – of the subfamily Colubrinae of the family Colubridae. They are medium to large constrictors and are found throughout much of the Northern Hemi ...
s and three species of
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most ...
s. One species of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. As of 2019, this family contained over 600 described genera and over 6,000 described species, making it the largest family of spiders at 13% of all sp ...
, '' Pelegrina tillandsiae'', has been found only on Spanish moss. Although widely presumed to infest Spanish moss, in one study of the ecology of the plant,
chiggers ''Trombicula'', known as chiggers, red bugs, scrub-itch mites, or berry bugs, are small arachnids (eight-legged arthropods) in the Trombiculidae family. In their larval stage, they attach to various animals, including humans, and feed on skin, ...
were not present among thousands of other arthropods identified on the plant.


Culture and folklore

Spanish moss often is associated with
Southern Gothic Southern Gothic is an artistic subgenre of fiction, country music, film and television that are heavily influenced by Gothic elements and the American South. Common themes of Southern Gothic include storytelling of deeply flawed, disturbing or ...
imagery and
Deep South The Deep South or the Lower South is a cultural and geographic subregion in the Southern United States. The term was first used to describe the states most dependent on plantations and slavery prior to the American Civil War. Following the war ...
culture, due to its propensity for growing in subtropical humid southern locales such as
Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...
,
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
,
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
,
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
,
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
,
east East or Orient is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fa ...
and
south South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
Texas, and extreme southern Virginia. One anecdote about the origin of Spanish moss is called "The Meanest Man Who Ever Lived", in which the man's white hair grew very long and got caught on trees. Spanish moss was introduced to Hawaii in the nineteenth century. It became a popular ornamental and lei plant. In Hawaii, it was named "ʻumiʻumi-o-Dole" after the beard of
Sanford B. Dole Sanford Ballard Dole (April 23, 1844 – June 9, 1926) was a lawyer and jurist from the Hawaiian Islands. He lived through the periods when Hawaii was a kingdom, protectorate, republic, and territory. A descendant of the American missionary ...
, the first president of the
Provisional Government of Hawaii The Provisional Government of Hawaii (abbr.: P.G.; Hawaiian: ''Aupuni Kūikawā o Hawaiʻi'') was proclaimed after the overthrow of the Hawaiian Kingdom on January 17, 1893, by the 13-member Committee of Safety under the leadership of its ch ...
. It also is known as hinahina, ("silvery") borrowing the name of the native heliotrope used in lei until shoreline development made access difficult. It has become a substitute for the native hinahina in lei used for pageantry. In the early 21st Century the plant was heavily marketed as " Pele's hair"/"lauoho-o-Pele," which actually refers to a type of filamentous
volcanic glass Volcanic glass is the amorphous (uncrystallized) product of rapidly cooling magma. Like all types of glass, it is a state of matter intermediate between the closely packed, highly ordered array of a crystal and the highly disordered array of liqu ...
.)


Human uses

Spanish moss has been used for various purposes, including building insulation, mulch, packing material, mattress stuffing, and fiber. In the early 1900s it was used commercially in the padding of car seats. More than 10,000 tons of processed Spanish-moss was produced in 1939. Today, it is collected in smaller quantities for use in arts and crafts, as bedding for flower gardens, and as an ingredient in
bousillage Bousillage (bouzillage,McDermott, John Francis. "bousillage, bouzillage, n. m.". ''A Glossary of Mississippi Valley French, 1673-1850''. St. Louis: , 1941. 34. Print. bousille, bouzille) is a mixture of clay and grass or other fibrous substances us ...
, a traditional wall covering material. In some parts of Latin America and Louisiana, it is used in Nativity scenes. In the desert regions of southwestern United States, sometimes dried Spanish moss is used in the manufacture of
evaporative cooler An evaporative cooler (also known as evaporative air conditioner, swamp cooler, swamp box, desert cooler and wet air cooler) is a device that cools air through the evaporation of water. Evaporative cooling differs from other air conditioning s ...
s, colloquially known as "swamp coolers" (and in some areas as "desert coolers"), which are used to cool homes and offices much less expensively than air conditioners. The cooling technology uses a pump that squirts water onto a pad made of Spanish moss plants; a fan then pulls air through the pad, and into the building. Evaporation of the water on the pads serves to reduce air temperature, cooling the building.


Varieties and cultivars

* ''Tillandsia'' 'Maurice's Robusta' * ''Tillandsia'' 'Munro's Filiformis' – a natural
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
with very fine, green leaves that is native to
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
and that also is known in the United States by the trade designations ''Tillandsia usneoides'' and , it conforms to the description of the now-defunct variety ''Tillandsia usneoides'' var. ''filiformis'' (
André André — sometimes transliterated as Andre — is the French and Portuguese form of the name Andrew, and is now also used in the English-speaking world. It used in France, Quebec, Canada and other French-speaking countries. It is a variation ...
) Mez * ''Tillandsia'' 'Odin's Genuina' : a natural variety with brown rather than green or yellow flower
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s that is native to
Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...
and
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
* ''Tillandsia'' 'Spanish Gold' * ''Tillandsia'' 'Tight and Curly'


Hybrids

* ''Tillandsia'' 'Nezley' (''Tillandsia usneoides'' × '' mallemontii'') * ''Tillandsia'' 'Kimberly' (''Tillandsia usneoides'' × '' recurvata'') * ''Tillandsia'' 'Old Man's Gold' ('' Tillandsia crocata'' × ''usneoides'')Bromeliad Cultivar Registry: Tillandsia 'Old Man's Gold'
/ref>


See also

* Lace lichen, an organism of similar habit and appearance


References

* Mabberley, D.J. 1987. ''The Plant Book. A Portable Dictionary of the Higher Plants''. Cambridge University Press, Cambridge. .


External links


Spanish Moss: Its History, Nature and Uses — Beaufort County Library




* ttps://www.leiday.org/whats-in-a-name-itla-okla-%ca%bb-umi%ca%bb-umi-o-dole-spanish-moss/ What’s in a name? Itla-okla, ʻUmiʻumi-o-Dole, Spanish Moss
Flowers Used for Lei
{{Taxonbar, from=Q311524 Tillandsia Plants described in 1762 Epiphytes Flora of the Caribbean Flora of Central America Flora of Mexico Flora of South America Flora of the Southeastern United States Flora of the South-Central United States Flora of the Bahamas Flora of Bermuda Flora without expected TNC conservation status